which

1. Does the first "which" refer to "images" or "circuits of neurons"?

2. Does "between which" refer to "between circuits of neurons" or "between circuits of neurons and bodily activities"?

Context:

The neuroscientist Antonio Damasio explains that one of the brain’s main tasks is continually to monitor and regulate our bodily functions, and it does this through establishing self-reflexive maps – images linking circuits of neurons which are established in the brain to correspond to different bodily activities, between which there is two-way feedback. This is essential to our well-being and obviously applies to routine physical functions such as the monitoring of body temperature, or the release of hormones to increase appetite stimulated by a fragrant waft of cooking, for example.