In turn

Salaam

Which meaning is more accurate for "in turn" in the following paragraph pleas?

1. Consequently

2. Then

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"At the time, neuroscientists had yet to figure out the mechanisms within the brain that allow animals to learn in this way. But Minsky was still able to loosely mimic the behavior, thereby advancing artificial intelligence. Several decades later, as reinforcement learning continued to mature, it in turn helped the field of neuroscience discover those mechanisms, feeding into a virtuous cycle of advancement between the two fields." (Source)