(a) An elephant was the heaviest animals on land. No. The elephant is one of the heaviest (of all the) land animals. (They are not extinct!) (b) The elephant is the land animals. No. The elephant is a land animal. (c) The elephant is living / lives on the land. OK
"The elephant" here refers to a particular species of animal, not to a particular individual.
Example:
The elephant is a heavy land animal that has a long probiscus and lives on land. (We are describing the characteristics of the animals that belong to the elephant species.)
Elephants (now we refer to the collection of all the living individuals)